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The first recorded case of a performing actor occurred in 534 BC (though the changes in calendar over the years make it hard to determine exactly) when the Greek performer Thespis stepped on to the stage at the Theatre Dionysus to become the first known person to speak words as a character in a play or story.
A lot of people claim it to be Thespis.
http://www.pbs.org/empires/thegreeks/background/24a_p1.html
According to legend, the first actor was an ancient Greek named Thespis. This is why actors are sometimes known as "Thespians".
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