it means he is either nervous or unsure, angry, bored or it may even just be a simple habit.
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Biting one's knuckle can be a sign of nervousness, excitement, or arousal. It may indicate that the person is feeling overwhelmed or trying to contain their emotions.
He's hungry
It means he realised he was too "forward" by smiling at you, so is now trying to control himself.
When a man bites my neck, it's an instant turn on. I mean, right now, find a bed. It took me a few years to develop this, as a young woman, I didn't like it. The older I got, the more I enjoyed biting, and being bitten.
You have startled the dog..scared him
A ant that bites a human that turns the human to a zombie