well depending on what skulls you turn on the skulls will make it harder [usually] or more fun [only two actually make it fun] but will improve your skills and give you more points
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I don't believe you collect any. You just turn them on or off.
In the wave options when customizing a firefight game.
When you have the skull menu brought up, click on the skulls you want to activate to activate them. Simply, move the control stick to the skull, and press the button.
it ether affects the gme to make it harder or it makes more fun like the grunt birtday party skull when you shoot a grunt in the head but all but two of the skulls make it harder so if you search this then you must like halo i would be suprised i you dont it is a great game if you want to know more then just ask me ok i know most there is to know about halo.
Red Skulls real name is Yo Hon Schmitz