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It is used to emulate an original 3-button gamepad for Genesis, because some games may glitch when you use a 6-button controller. To emulate an original 3-button controller, you should insert a game cartridge into Genesis console and hold the mode button before turning it on. Then turn it on, wait while the game boots up, and release the mode button. Now your controller should work as 3-button one. For example, in Mortal Kombat II there are 5 actions you can do: Low Punch, High Punch, Low Kick, High Kick and Block. If you have a 6-button controller then you have enough buttons for this game. For those, who don't have a 6-button controller, MKII supports 3-button gamepads too! Simply press A and B buttons simultaneously to perform High Punch or B and C buttons to perform High Kick (default button layout is used). This trick only works when using a 3-button controller, or 6-button controller in emulation mode. Also, the mode button does something in some games, but there are only few of them.

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Q: What is the use of the mode button on the Sega Genesis?
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