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Yes United States Nintendo DS games can be played on a Nintendo DS UK console. This is because DS games are not locked according to PAL regions.
An R4 is basically a mod chip for the Nintendo DS console. You can download games from the internet and put them on your R4 card and play the games on your Nintendo DS.
Yes, the original Nintendo DS as well as the Nintendo DS lite can play NDSi games.
You download them off the Wii shop channel, which lets you play them on the Virtual Console.
No. There is no way to download or play DSiWare games on Nintendo DS or DS Lite systems.