Yes, Moro (the eldest wolf and San's adopted mother) dies, but the pups do not
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No , Ashitaka and San return the head and all becomes green again - new life .
Spoiler Alert!No, none of the wolves die.
Princess Tenagnework died on 2003-04-06.
Kurun Princess Hexiao died on 1823-10-13.
Princess Soraya died on October 25, 2001, in Paris, France of brain hemorrhage.